The purpose of the crossovers is to limit the frequency range being played by the speakers. In your case, you would specifically want to limit the amount of low frequency content being reproduced by both your front and rear speakers. If you don't have a crossover you will be using to highpass both sets of speakers, such as one that may be built into the headunit, then you will need to use the crossovers built into the amps.
The PX300.4 offers the nice feature of selectable 12db/oct or 24db/oct slopes. This is a feature the GTO75.4II does not offer.
But if your headunit has built in crossovers, this may be a feature you won't be using on the amplifier.